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・Kyoto University

・Institute for Frontier Life and Medical Sciences

・Graduate School of Medicine and Faculty of Medicine

・Graduate School of Biostudies

・ Institute for Integrated Cell-Material Sciences（iCeMS）

・Scientific Research on Innovative Areas「Interplay of developmental clock and extracellular environment in brain formation」

・ Japan Science and Technology Agency(CREST)　[Opt Bio] Development and application of optical technology for spatiotemporal control of biological functions

・Japan Agency for Medical Research and Development (AMED-CREST)「成体脳神経幹細胞の機能低下メカニズムの解明と再活性化に向けた技術開発」